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{ "pk": 47113, "title": "Paleogene marine bivalves of the deep-water Keasey Formation in Oregon, Part IA: The protobranchs (Nuculidae, Sareptidae, Pristiglomidae)", "subtitle": null, "abstract": "<p>Systematic description and illustration of protobranch bivalves of the late Eocene–early Oligocene Keasey Formation in Oregon follows previous monographic treatments of the Keasey anomalodesmatans, heteroconchs and pteriomorphs. Of the three protobranch families documented here, this is the first recognition of Sareptidae and Pristiglomidae in the fossil record of the Northeastern Pacific and Cascadia Margin, as well as recognition of three undescribed species of Nuculidae. Small size and poor preservation contribute to previous failure to notice them in the field. New taxa include <em>Nucula</em> (<em>Nucula</em>) <em>cascadensis</em>, <em>Ennucula keaseyesis</em>, <em>Sarepta oregonensis</em>,<em> Sarepta nascif</em>, and <em>Pristigloma mistensis</em>. <em>Nucula</em> (<em>Nucula</em>) n. sp. is treated in open nomenclature pending discovery of additional material.<em> Acila</em> (<em>Truncacila</em>) <em>nehalemensis</em> is one of the most abundant and frequent bivalves in the formation. Because of the longstanding importance of the truncate acilas in Paleogene biostratigraphy, comparative accounts and illustrations are provided for three superposionally important species from the Eugene and Pittsburg Bluff formations in Oregon and the Twin River Formation in Washington. The Keasey Protobranch Association is an important component of the transitional peri-seep biotope at three distinctive Keasey methane seeps. The association also occurs throughout the formation at localities where protobranchs are preserved in fine-grained tuffaceous siltstones with gadilinid scaphopods and deposit-feeding taxa with living relatives adapted to dysoxia, diffuse methane seepage, and toxic geochemistry. Treatments include global biogeographic references and comparisons with similar taxa and molluscan assemblages, notably in the basins of southern New South Wales, South Australia, Western Australia, New Zealand, and Antarctica as well as active Western Pacific Margin settings in northern Japan and Far Eastern Russian Kuril Islands, Sakhalin Island, and Kamchatka Peninsula</p>", "language": "eng", "license": { "name": "Creative Commons Attribution-NonCommercial-ShareAlike 4.0", "short_name": "CC BY-NC-SA 4.0", "text": "<p><!-- x-tinymce/html --></p>\n<p>Readers are free to:</p>\n<ul>\n<li><strong>Share</strong> — copy and redistribute the material in any medium or format</li>\n<li><strong>Adapt</strong> — remix, transform, and build upon the material<br><br>The licensor cannot revoke these freedoms as long as you follow the license terms.</li>\n</ul>\n<p>Under the following terms:</p>\n<ul>\n<li><strong>Attribution</strong> — You must give appropriate credit , provide a link to the license, and indicate if changes were made . You may do so in any reasonable manner, but not in any way that suggests the licensor endorses you or your use.</li>\n<li><strong>NonCommercial</strong> — You may not use the material for commercial purposes .</li>\n<li><strong>ShareAlike</strong> — If you remix, transform, or build upon the material, you must distribute your contributions under the same license as the original.<br><br>No additional restrictions — You may not apply legal terms or technological measures that legally restrict others from doing anything the license permits.</li>\n</ul>\n<p>Notices:</p>\n<p>You do not have to comply with the license for elements of the material in the public domain or where your use is permitted by an applicable exception or limitation.</p>\n<p>No warranties are given. The license may not give you all of the permissions necessary for your intended use. For example, other rights such as publicity, privacy, or moral rights may limit how you use the material.</p>", "url": "https://creativecommons.org/licenses/by-nc-sa/4.0" }, "keywords": [ { "word": "taxodont dentition" }, { "word": "nacre" }, { "word": "Eocene-Oligocene transition" }, { "word": "Cascadia Margin" }, { "word": "methane seep" }, { "word": "peri-seep biotope" } ], "section": "Article", "is_remote": true, "remote_url": "https://escholarship.org/uc/item/3c46d2rh", "frozenauthors": [ { "first_name": "Carole", "middle_name": "S.", "last_name": "Hickman", "name_suffix": "", "institution": "UC Berkeley", "department": "" } ], "date_submitted": "2025-03-31T23:40:26.338000Z", "date_accepted": "2025-07-08T22:05:03.548000Z", "date_published": "2025-07-10T23:10:00Z", "render_galley": { "label": "PDF", "type": "pdf", "path": "https://journalpub.escholarship.org/ucmp_paleobios/article/47113/galley/36867/download/" }, "galleys": [ { "label": "PDF", "type": "pdf", "path": "https://journalpub.escholarship.org/ucmp_paleobios/article/47113/galley/36867/download/" } ] }